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A calibrated mizerReef object for a generic Caribbean coral reef with ten functional groups, parameterised from field data collected at Karpata Reef, Bonaire (FORCE dataset, Dryden 2017). Functional groups span six predator guilds, parrotfish, farming damselfish, other herbivores, and invertebrates. The model includes competitive predation refuge (using karpata_refuge) and benthic resources (algae and detritus).

Usage

caribbean_10_model

Format

A mizerReef object (extends MizerParams) with 10 species, competitive refuge, and algae/detritus components.

Source

PhD Thesis, FORCE dataset

Details

Built from caribbean_10_species, caribbean_10_interaction, and karpata_refuge; calibrated to match observed biomasses and built with mizerReef 2.0.0 / mizer 3.1.0. Detritus resource dynamics are tuned via tuneUR() so that they are genuinely at steady state (dB/dt = 0) for the model's calibrated abundances. Algae biomass is likewise tuned by tuneUR() to its own steady state, but for a fixed, literature-informed production rate (algae_growth, left at newReefParams()'s default of 2000 grams per square meter per year – see setAlgaeParams) that is not itself retuned to match consumption, since real algal production is not driven by grazer demand; an earlier untuned version of this object (algae biomass not yet solved for its steady state) is archived at inst/archive/caribbean_10_model_untuned.rda for reference.

Recalibrated in 2026 against a corrected senescence-mortality formula (reefSenMort()), the same underlying issue that required recalibrating caribbean_3_model. Biomass, growth, and reproduction were all re-tuned (verified: repeated reefSteady() calls leave biomass unchanged). Final biomass is within 79-121\ six FORCE-observed targets (pred_eng, pred_grab, pred_inv, pred_plank, parrotfish, herbs); growth (age at maturity) matches its target closely for five of those six, and reasonably (within a factor of ~3) for pred_grab after correcting its own age_mat target (see caribbean_10_species's documentation). eels and farm_damsel – which have no FORCE-observed biomass target – were given soft, literature-derived targets and brought up from near-zero to a modest, non-vanishing presence, deliberately not matched exactly (doing so costs disproportionate accuracy on the six well-observed groups). A version of this model that ignores eels/farm_damsel entirely and matches the six FORCE-observed groups more tightly is archived (not bundled as a package dataset, to avoid doubling the recalibration burden) at inst/archive/caribbean_10_model_tight_biomass_fit.rds – load with mizer::readParams(). The complexity-increases-biomass relationship central to the associated manuscript/thesis chapter was explicitly re-verified against this recalibrated model (competitive vs. non-complex refuge), not merely assumed to still hold. See inst/scripts/Caribbean_10_model-calibration.R's design note for the full reasoning, including why growth-matching required tuning one species at a time rather than mizerExperimental's usual batched workflow.

References

FORCE dataset. From: Dryden, C. (2017). Habitat structural complexity of Caribbean coral reefs and its relationships with fish community structure. PhD Thesis, Newcastle University.

Beese, C. (2025). PhD Thesis. Victoria University of Wellington. https://doi.org/10.26686/wgtn.26421523